- Johnston SE, Beraldi D, McRae AF, Pemberton JM, Slate J (2010). Horn type and horn length genes map to the same chromosomal region in Soay sheep. Heredity,104(2): 196-205. [see all QTL
 by this paper]
- Johnston SE, McEwan J, Pickering N, Kijas J, Beraldi , Pilkington J, Pemberton J, Slate J (2011). Genome-wide association mapping identifies the genetic basis of discrete and quantitative variation in sexual weaponry in a wild sheep population. Molecular ecology,20(12):2555-66. [see all QTL by this paper]
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